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Scientific and other Publications from 1998/99 to 2018

Since 1997 the Foundation supports Scientific Research Projects with the emphasis on shark preservation. The results of some of the research projects have been published in well known scinetific journals, as posters or in other forms.


  > Identification (15)
> Fiji Migration Project (7)
> Fiji Shark Reef Marine Reserve (4)
> Capture Stress (2)
> Catch and Release (2)
> Nurseries (1 + Poster)
> Basking shark project (2)
> Lemon Sharks in Jupiter, FL, USA (1 + 1 conference paper)
> Whale sharks (11)
> Angel Sharks Gran Canaria (2 Poster)
> Behaviour (1)
> Masters thesis on shovelnose hammerhead sharks



D. Pinhal, M. S. Shivji, M. Vallinoto, D. D. Chapman, O. B. F. Gadig and C. Martins   2012
Cryptic hammerhead shark lineage occurrence in the western South Atlantic revealed by DNA analysis
Marine Biology159:829-836 (2012)
> Download PDF (762 kB)

D. D. Chapman, D. Pinhal, M. S. Shivji   2009
Tracking the fin trade: genetic stock identification in western Atlantic scalloped hammerhead sharks Sphyrna lewini
Endangered Species Research Vol. 9: 221 - 228

D. D. Chapman, B. Fichau and M. S. Shivji   2008
Parthenogenesis in a large-bodied requiem shark, the blacktip Carcharhinus limbatus
Journal of Fish Biology 73, 1473-1477

H. Sebastian, P. A. Haye and M. S. Shivji   2008
Characterization of the pelagic shark-fin trade in north-central Chile by genetic identification and trader surveys
Journal of Fish Biology 73, 2293-2304

J. E. Magnussen, E. K. Pikitch, S. C. Clarke, C. Nicholson, A. R. Hoelzel & M. S. Shivji   2007
Genetic tracking of basking shark products in international trade
Animal Conservation 10 199-207.

Shelley C. Clarke, Murdoch K. McAllister, E. J. Milner-Gulland, G. P. Kirkwood, Catherine G. J. Michielsens, David J. Agnew, Ellen K. Pikitch, Hideki Nakano and Mahmood S. Shivji   2006
Global estimates of shark catches using trade records from commercial markets
Ecology Letters 9: 1115 1126

Shelley C. Clarke, Jennifer E. Magnussen, Debra L. Abercombie, Murdoch K. McAlister and Mahmood S. Shivji   2005
Identification of Shark Species Composition and Proportion in the Hong Kong Shark Fin Market Based on Molecular Genetics and Trade Records
Conservation Biology Volume 20, No. 1, 201-211

Debra L. Abercombie, Shelley C. Clarke Mahmood S. Shivji   2005
Global-scale genetic identification of hammerhead sharks: Application to assessment of the international fin trade and law enforcement
Conservation Genetics 6:775-788

Mahmood S. Shivji, Demian D. Chapman, Ellen K. Pikitch & Paul W. Raymond   2005
Genetic profiling reveals illegal international trade in fins of the great white shark, Carcharodon carcharias
Conservation Genetics 6:1035-1039

Chapman, D.D., P.A. Prodohl, J. Gelsleichter, C.A. Manire and M.S. Shivji   2004
Predominance of genetic monogamy by females in a hammerhead shark, Sphyrna tiburo: Implications for shark conservation.
Molecular Ecology 13: 1965-1974.

C. J. Douady, M. Dosay, M. S. Shivji, and M. J. Stanhope   2003
Molecular phylogenetic evidence refuting the hypothesis of Batoidea (rays and skates) as derived sharks.
Molecular Phylogenetics and Evolution 26: 215-221.

Pank, M., L. Natanson, N. Kohler, M. Stanhope and M. Shivji   2003
Rapid species identification of pelagic shark tissues using genetic approaches.
In: Sharks of the Open Ocean. E. Pikitch & M. Camhi (Eds.). Blackwell Science Inc. (In Press).

Demian D. Chapman, Debra L. Abercombie, Christophe J. Douady, Ellen K. Pikich, Michael J. Stanhope and Mahmood S. Shivji   2003
A streamlined, bi-organelle, multiplex PCR approach to species identification: Application to global conservation and trade monitoring of the greatwhite shark, Carcharodon carcharias
Conservation Genetics 4: 415-425

M. Shivji, S. Clarke, M. Pank, L. Natanson, N. Kohler and M. Stanhope   2002
Genetic Identification of Pelagic Shark Body Parts for Conservation and Trade-Monitoring.
Conservation Biology, 16(4):1036-1047.

M. Pank, M. Stanhope, L. Natanson, N. Kohler and M. Shivji   2001
Rapid and simultaneous identification of body parts from the morphologically similar sharks Carcharhinus obscurus and Carcharhinus plumbeus (Carcharhinidae) using multiplex PCR.
Marine Biotechnology 3:231-240.

Fiji Migration Project


Juerg M. Brunnschweiler, Adam Barnett   2013
Opportunistic Visitors: Long-Term Behavioural Response of Bull Sharks to Food Provisioning in Fiji
PLoS ONE 8(3): e58522. doi:10.1371/journal.pone.0058522

C. Testerman, J. Brunnschweiler, M. Heithaus, S. Gulak, J. Werry, R. Jabado, C. Jones and M. Shivji   2012
Global Population Genetic Structure and Parentage Analysis of the Bull Shark (Carcharhinus leucas)
AES Conference 2012 Poster
> Download Poster PDF (299 kB)

Brunnschweiler J. M., Harld Baensch   2010
Seasonal and Long-Term Changes in Relative Abundance of Bull Sharks from a Tourist Shark Feeding Site in Fiji
PLoS ONE 6(1): e16597. doi:10.1371/journal.pone.0016597

Brunnschweiler J. M., N. Queiroz and W. Sims   2010
Oceans apart? Short-term movements and behaviour of adult bull sharks Carcharhinus leucas in Atlantic and Pacific Oceans determined from pop-off satellite archival tagging
Journal of Fish Biology doi:10.1111/j.1095-8649.2010.02757.x

Brunnschweiler J. M.   2009
Tracking free-ranging sharks with hand-fed intra-gastric acoustic transmitters
Marine and Freshwater Behaviour and Physiology (2009) Vol. 42, No. 3, May 2009, 201-209
> Download PDF (730 kB)

Brunnschweiler J. M.   2008
Species record and mistaken identifications: the case of bull sharks at Chumphon Pinnacle, Thailand.
JMBA2 - Biodiversity Records: published online

Brunnschweiler J. M., Compagno L.J.V.   2007
First record of Carcharhinus leucas from Tonga, South Pacific.
JMBA2 - Biodiversity Records: published online

Juerg M. Brunnschweiler & John L. Earle   2006
A contribution to marine life conservation efforts in the South Pacific: The Shark Reef Marine Reserve, Fiji
Cybium 30(4) suppl.: 133-139.

Fiji Shark Reef Marine Reserve


Brunnschweiler J. M. 2009
The Shark Reef Marine Reserve: a marine tourism project in Fiji involving local communities
Journal of Sustainable Tourism 99999:1
> Download PDF (811 kB)

Brunnschweiler J.M.   2008
An integrated approach to study bull shark behaviour and ecology in the South Pacific: The Bull Shark Tagging Programme.
12th annual meeting of the European Elasmobranch Association, Lisbon.

Brunnschweiler J.M., Earle, J.L.   2006
A contribution to marine life conservation efforts in the South Pacific: The Shark Reef Marine Reserve, Fiji.
Cybium 30 (Supplement): 133-139

Brunnschweiler J.M., Sazima I.   2006
A new and unexpected host for the sharksucker (Echeneis naucrates) with a brief review of the echeneid-host interactions.
JMBA2 - Biodiversity Records: published online

Capture Stress


Hyatt, M., Anderson, P., O'Donnell, P.   2013
After Capture and Handling, Behavioral Assessment of Bull (Carcharhinus leucas) and Bonnethead (Sphyrna tiburo) Sharks at Release is a Poor Indicator of Physiological Stress. 38th Annual Eastern Fish Health Workshop. Gettysburg, PA.

Michael W. Hyatt, Paul A. Anderson, Patrick M. O'Donnell, Ilze K. Berzins   2012
Assessment of acid-base derangements among bonnethead (Sphyrna tiburo), bull (Carcharhinus leucas), and lemon (Negaprion brevirostris) sharks from gillnet and longline capture and handling methods
Comparative Biochemistry and Physiology, Part A
> Download PDF (590 kB)

Catch and Release


A. J. Gallagher, P. M. Kyne and N. Hammerschlag   2012
Ecological risk assessment and its application to elasmobranch conservation and management
Journal of Fish Biology 1095-8649.2012
> Download PDF (688 kB)

Neil Hammerschlag and James Sulikowski   2011
Killing for conservation: the need for alternatives to lethal sampling of apex predatory sharks
Endangered Species Research Vol. 14: 135-140, 2011
> Download PDF (119 kB)



Steiner, PA, Michel, M, O'Donnell, PM   2007
Notes on the Occurrence and Distribution of Elasmobranchs in the Ten Thousand Islands Estuary, Florida. Pages 237-250 in C.T. McCandless, N.E. Kohler, and H.L. Pratt, Jr., editors. Shark nursery grounds of the Gulf of Mexico and the east coast waters of the United States. American Fisheries Society, Symposium 50, Bethesda, Maryland.

Presentation of the first results of the Rookery Bay / Tenthousand Islands shark nursery project by Pat O'Donnel:
> Rookery Bay Poster (PDF, 304 kb)

Basking shark project


Matthew J. Witt, Tom Hardy, Louise Johnson, Catherine M. McClellan, Stephen K. Pikesley, Sue Ranger, Peter B. Richardson, Jean-Luc Solandt, Colin Speedie, Ruth Williams and Brendan J. Godley   2012
Basking sharks in the northeast Atlantic: spatio-temporal trends from sightings in UK waters
Marine Ecology Progress Series Vol. 459: 121-134, 2012; doi: 10.3354/meps09737
> Download PDF (1.1 MB)

E.J. Southall, D.W. Sims, J.D. Metcalfe, J.I. Doyle, S. Fanshawe, C. Lacey, J. Shrimpton, J.-L. Solandt and C.D. Speedie   2005
Spatial distribution patterns of basking sharks on the European shelf: preliminary comparison of satellite-tag geolocation, survey and public sightings data.
J. Mar. Biol. Ass. U.K. 85, 1083-1088

Lemon Sharks in Jupiter, FL, USA


Conference Paper:
J. Baeyaert, F. Dhellemmes, K. Erzini, S. H Gruber and T. L. Guttridge  2016
Wild Spatial Behaviour & Personality Traits: A comparison study for juvenile lemon sharks
Rapp. Comm. int. Mer Médit., 41, 2016

Kessel S. T., Chapman D. D., Franks B. R., Gedamke T., Gruber S. H., Newman J. M., White E. R. and Perkins R. G.   2014
Predictable temperature regulated residency, movement and migration in a large, highly-mobile marine predator
Marine Ecology Progress Series 514:175-190 November 2014
doi: 10.3354/meps10966

Whale sharks


Andrzejaczek S, Meeuwig J, Rowat D, Pierce S, Davies T, Fisher R, Meekan M.  2016
The ecological connectivity of whale shark aggregations in the Indian Ocean: a photo-identification approach.
R. Soc. open sci.3
doi: 10.1098/rsos.160455

Christoph A. Rohner, Anthony J. Richardson, Clare E.M. Prebble, Andrea D. Marshall, Michael B. Bennett, Scarla J.Weeks, Geremy Cliff, Sabine P.Wintner and Simon J. Pierce   2015
Laser photogrammetry improves size and demographic estimates for whale sharks.
PeerJ 3:e886
doi: 10.7717/peerj.886

Cagua EF, Cochran JEM, Rohner CA, Prebble CEM, Sinclair-Taylor TH, Pierce SJ, Berumen ML.   2015
Acoustic telemetry reveals cryptic residency of whale sharks.
Biol. Lett. 11: 20150092.

Thomas M. Vignaud, Jeffrey A. Maynard, Raphael Leblois, Mark G. Meekan, Ricardo Vazquez-Juarez, Deni Ramirez-MacIas, Simon J. Pierce, David Rowat, Michael L. Berumen, Champak Beeravolu, Sandra Baksay and Serge Planes   2014
Genetic structure of populations of whale sharks among ocean basins and evidence for their historic rise and recent decline
Molecular Ecology (2014) 23, 2590-2601
doi: 10.1111/mec.12754

Christoph A. Rohner, Lydie I. E. Couturier, Anthony J. Richardson, Simon J. Pierce, Clare E. M. Prebble, Mark J. Gibbons, Peter D. Nichols   2013
Diet of whale sharks Rhincodon typus inferred from stomach content and signature fatty acid analyses
doi: 10.3354/meps10500

S. Fox, I. Foisy, R. De La Parra Venegas, B. E. Galvan Pastoriza, R. T. Graham, E. R. Hoffmayer, J. Holmberg and S. J. Pierce   2013
Population structure and residency of whale sharks Rhincodon typus at Utila, Bay Islands, Honduras
Journal of Fish Biology
doi: 10.1111/jfb.12195

C. A. Rohner, S. J. Pierce, A. D. Marshall, S. J. Weeks, M. B. Bennett, A. J. Richardson   2013
Trends in sightings and environmental influences on a coastal aggregation of manta rays and whale sharks
doi: 10.3354/meps10290

David P. Robinson, Mohammed Y. Jaidah, Rima W. Jabado, Katie Lee-Brooks, Nehad M. Nour El- Din, Ameena A. Al Malki, Khaled Elmeer, Paul A. McCormick, Aaron C. Henderson, Simon J. Pierce, Rupert F. G. Ormond   2013
PLOS ONE ( March 2013, Vol 8, Issue 3 | e58255

Juerg M. Brunnschweiler and David W. Sims   2011
Diel Oscillations in Whale Shark Vertical Movements Associated with Meso- and Bathypelagic Diving
American Fisheries Society Symposium 76, 2011
> Download PDF (1.4 MB)

N. E. Humphries, N. Queiroz, J. R. M. Dyer, N. G. Pade, M. K. Musyl, K. M. Schaefer, D. W. Fuller, J. M. Brunnschweiler, T. K. Doyle, J. D. R. Houghton, G. C. Hays, C. S. Jones, L. R. Noble, V. J. Wearmouth, E. J. Southall & D. W. Sims   2010
Environmental context explains Lévy and Brownian movement patterns of marine predators
Nature doi:10.1038/nature09116

Brunnschweiler J.M., Baensch H., Pierce S.J., Sims D.W.   2009
Deep-diving behaviour of a whale shark Rhincodon typus during long-distance movement in the western Indian Ocean.
Journal of Fish Biology 74: 706-714

Angel Sharks Gran Canaria


Poster presentation of Gran Canarian Angel Shark "Stomach Contents Analysis" at the 15th Spanish symposium of marine biological research, 19th Sep. 2008.
> Poster (JPG, 265k) Spanish

Poster presentation at "Davy Jones Diving" on the results of the Angel Shark survey from May 2006 to August 2008 in Gran Canaria.
> Poster (JPG, 343k) Spanish/English




Ritter, Erich K. and Godknecht, Alexander J.   2000
Agonistic displays in the Blacktip Shark (Carcharhinus limbatus).
Copeia February 1, 2000. 282-284.

Masters thesis on shovelnose hammerhead sharks


Influences of biogeographic changes in the Gulf of Mexico on molecular population structures of the shovelnose hammerhead-shark Sphyrna tiburo.
Markus Ruch, Masters thesis at the Zoological Museum, University of Zürich, January 2001.

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